TY - GEN AB - Gene expression noise is an evolvable property of biological systems that describes differences in expression among genetically identical cells in the same environment. Prior work has shown that expression noise is heritable and can be shaped by selection, but the impact of variation in expression noise on organismal fitness has proven difficult to measure. Here, we quantify the fitness effects of altering expression noise for the TDH3 gene in Saccharomyces cerevisiae. We show that increases in expression noise can be deleterious or beneficial depending on the difference between the average expression level of a genotype and the expression level maximizing fitness. We also show that a simple model relating single-cell expression levels to population growth produces patterns consistent with our empirical data. We use this model to explore a broad range of average expression levels and expression noise, providing additional insight into the fitness effects of variation in expression noise. AD - University of Michigan AD - University of Michigan AD - University of Chicago AD - University of Michigan AD - University of Michigan AD - University of Michigan AD - University of Michigan AD - University of Michigan AU - Duveau, Fabien AU - Hodgins-Davis, Andrea AU - Metzger, Brian P. H. AU - Yang, Bing AU - Tryban, Stephen AU - Walker, Elizabeth A. AU - Lybrook, Tricia AU - Wittkopp, Patricia J. DA - 2018-08-20 ID - 9902 JF - eLife KW - gene expression KW - promoter KW - TDH3 KW - evolution KW - selection KW - competitive growth L1 - https://knowledge.uchicago.edu/record/9902/files/elife-37272-v2.pdf L1 - https://knowledge.uchicago.edu/record/9902/files/elife-37272.zip L2 - https://knowledge.uchicago.edu/record/9902/files/elife-37272-v2.pdf L2 - https://knowledge.uchicago.edu/record/9902/files/elife-37272.zip L4 - https://knowledge.uchicago.edu/record/9902/files/elife-37272-v2.pdf L4 - https://knowledge.uchicago.edu/record/9902/files/elife-37272.zip LA - eng LK - https://knowledge.uchicago.edu/record/9902/files/elife-37272-v2.pdf LK - https://knowledge.uchicago.edu/record/9902/files/elife-37272.zip N2 - Gene expression noise is an evolvable property of biological systems that describes differences in expression among genetically identical cells in the same environment. Prior work has shown that expression noise is heritable and can be shaped by selection, but the impact of variation in expression noise on organismal fitness has proven difficult to measure. Here, we quantify the fitness effects of altering expression noise for the TDH3 gene in Saccharomyces cerevisiae. We show that increases in expression noise can be deleterious or beneficial depending on the difference between the average expression level of a genotype and the expression level maximizing fitness. We also show that a simple model relating single-cell expression levels to population growth produces patterns consistent with our empirical data. We use this model to explore a broad range of average expression levels and expression noise, providing additional insight into the fitness effects of variation in expression noise. PY - 2018-08-20 T1 - Fitness effects of altering gene expression noise in <i>Saccharomyces cerevisiae</i> TI - Fitness effects of altering gene expression noise in <i>Saccharomyces cerevisiae</i> UR - https://knowledge.uchicago.edu/record/9902/files/elife-37272-v2.pdf UR - https://knowledge.uchicago.edu/record/9902/files/elife-37272.zip Y1 - 2018-08-20 ER -